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A. Pentecost (born 1948)

Academic profile

Schools attended:
1952-55 Charles Street Primary, Southborough
1955-60 St Peters School, Southborough
1960-61 Ridgewaye School, Southborough
1961-67 Tunbridge Wells Technical School

Higher education and fellowships etc:
1967-71 Scholarship in physics and chemistry, Imperial College London
1967-68 Royal School of Mines, Imperial College, University of London
1968-1971 Royal College of Science. BSc Hons class 1. Botany (spec. plant physiology) and Associate of the Royal College of Science.
1971-75 University College of North Wales, Marine Science Laboratories PhD in algal ecophysiology under G.E. Fogg FRS.
1975-78 School of Plant Biology, University College of North Wales, Bangor.
1975-78 NERC Research Assistantship with Dr. C. Wood. Algal ecophysiology.
1978 Lectureship Chelsea College, University of London (later King's College London) specialising in algal ecology, ecoststem modelling
1998 Fellow of the Royal Statistical Society
1999 DSc in Geomicrobiology and algal ecophysiology, University of London.
2000 Readership in Geomicrobiology at King's College.
2007 Emeritus Reader Kings College London.
2007 Honorary Research Fellow of the Freshwater Biological Association.

Current research activities based mainly at the Freshwater Biological Association, Cumbria.

These include:
A review of the subaerial cyanobacteria (jointly with BA Whitton) exploring adaptations to water stress, high irradiance and temperature, analysing communities, nutrient relationships and inorganic carbon acquisition, colonisiation, endolithic activity and bio-erosion.

Changes in the fossil algal flora in postglacial marl deposits from Malham Tarn, Yorkshire, relating these to climate and human-induced catchment change over the past 8000 years.


Pentecost, A. 1968. The lichen-flora of the Lizard. Liz. Nat. 3, 4-11. 2. 

Pentecost, A. 1972. Imperial College Expedition to the Ruwenzori Mountains, Uganda. Explor.Rev. 2, 23-29.

Pentecost, A. & Fletcher, A. 1974. Tufa, (travertine) an unusual lichen substrate. Lichenologist 6, 100-102.

Pentecost, A. 1977.  A comparison of the flora of two mountain streams in Gwynedd. Lichenologist 9, 107-111.
Pentecost, A. & Happey-Wood, C.M. 1978. Primary production studies in two linked but contrasting Welsh lakes. Freshwat. Biol. 8, 9-24.

Pentecost, A. 1978. Blue-green algae and freshwater carbonate deposits. Proc. R. Soc. Lond. B. 200, 43-61.

Pentecost, A. l978. Calcification and photosynthesis in Corallina officinalis using the 14-C method. Br. Phycol. J. 13, 383-390.

Pentecost, A. 1979. Aspect and slope preferences in a saxicolous lichen community. Lichenologist 11, 81-83.

Pentecost, A. 1980. Aspects of competition in saxicolous lichen communities. Lichenologist 12, 135-144.

Pentecost, A. 1980. Calcification in plants.  Int. Rev. Cytol. 62, 1-27.

Pentecost, A.1980. The lichens and bryophytes of rhyolite and  pumice-tuff rock outcrops in Snowdonia, and some factors affecting their distribution. J. Ecol. 68, 251-67.

Pentecost, A. 1981. The tufa (travertine) deposits of the Malham district, North Yorkshire. Fld. Stud. 5, 365-87.

Happey-Wood, C. M. & Pentecost, A. 1981. Algal bioassay of  the water from two linked but contrasting Welsh lakes. Freshwat. Biol. 11, 473-491.
Pentecost, A. 1981. Some observations on the size and shape of  ascospores in relations to ecology and taxonomy. New Phytol. 89, 667-678.

Pentecost, A. 1982. A quantitative study of calcareous and  Tintenstriche algae from the Malham district, Northern  England. Br. Phycol. J. 17, 443-456.

Pentecost, A. 1982. The distribution of Euglena mutabilis in  Sphagna, with reference to Malham Tarn North Fen. Fld. Stud. 5, 591-606.

Pentecost, A. 1982. Conservation of freshwater algae. Ecos. 3, 8-11.
Pentecost, A. 1983. The distribution of daughter colonies in a  natural population of Volvox aureus Ehrenb.  Ann. Bot. 52, 769-776.
Pentecost, A. 1984. The growth of Chara globularis and its relationship to calcium carbonate deposition in Malham Tarn. Fld. Stud. 6, 53-58.
Livingstone, D., Pentecost, A. & Whitton, B.A. 1984. Diel variations in nitrogen and carbon dioxide fixation by  the blue-green alga Rivularia in an upland stream. Phycologia 23, 125-133.
Pentecost, A. 1984. Effects of sedimentation and light  intensity on mat forming Oscillatoriaceae with particular reference to Microcoleus lyngbyaceus Gomont. J. Gen. Microbiol . 130, 983-990. 

Pentecost, A. 1984. Introduction to Freshwater Algae. Richmond  Publishing Co., Surrey. 256 pp.

Pentecost, A. 1984. Observations on a bloom of the neuston alga  Nautococcus pyriformis from southern England with an explanation of the floatation mechanism. Br. Phycol. J. 19, 227-232. 

Pentecost, A. 1985. Photosynthetic plants as intermediary  agents between environmental bicarbonate and carbonate  deposition. In: (W.J. Lucas & J. A. Berry, eds.) Inorganic carbon uptake by aquatic photosynthetic plants.  Rockville, Maryland. pp. 459-476. 

Pentecost, A. &  Rose, C. I. 1985. Apothecium production in Xanthoria parietina. Lichenologist. 17, 33-38.

Pentecost, A. 1985. Investigation of variation in heterocyst  numbers, sheath development and false-branching in natural  populations of Scytonemataceae. Arch. Hydrobiol. 102, 343-353.

Pentecost, A. 1985. Association of cyanobacteria with tufa  deposits : identity, enumeration and nature of the sheath  material revealed by histochemistry. Geomicrobiol. J. 4,  285-298.

Pentecost, A. 1985. Alnus leaf impressions from a  postglacial tufa (travertine) in Yorkshire. Ann. Bot56, 779-782.

Pentecost, A. 1985.  Relationships between light, temperature and photosynthesis in a temperate Microcoleus (cyanobacterium) mat.  Microbios. 43, 141-148.

Pentecost, A. &  Rose, F. 1985. Changes in the cryptogam flora of the Wealden sandrocks, 1688-1984. Bot. J. Linn. Soc. 90, 217-230.

Pentecost, A. &  Riding, R. 1986. Calcification in Cyanobacteria. In: Biomineralization of Lower Plants and  Animals  (B.S.C. Leadbeater & R. Riding, eds.) pp. 73-90. Oxford.

Okazaki, M., Pentecost, A., Tanaka, Y. & Miyata, M. 1986. A study of calcium carbonate deposition in the genus Padina (Phaeophyceae, Dictyotales). Br. Phycol. J. 21, 217-224.

Pentecost, A. 1987. Some observations on the growth rates  of mosses associated with tufa (travertine) deposits and the interpretation of some Postglacial bryoliths. J. Bryol. 14, 543-550.

Pentecost, A. 1987. Growth and calcification of the freshwater cyanobacterium Rivularia haematitesProc. R. Soc. Lond. B. 232, 125-136.

Pentecost, A. 1987. The lichen-flora of Gwynedd. Lichenologist 19, 97-167.

Pentecost, A. & Bauld, J. 1988. Nucleation of calcite on the sheaths of cyanobacteria using a simple diffusion cell. Geomicrobiol J. 6, 129-135.

Pentecost, A. 1988. Estimates of sheath synthesis rates in a  motile cyanobacterium using a microautoradiographic method. Microbios 53, 109-117.  38

Pentecost, A. 1988. Observations on growth rates and calcium  carbonate deposition in the green alga Gongrosira. New. Phytol. 110, 249-253.

Pentecost, A. 1988. Growth and calcification of the cyanobacterium Homoeothrix crustacea. J. Gen. Microbiol. 134, 2665-2671.

Pentecost, A. & Lord, T. C. 1988. Postglacial tufas and  travertines from the Craven district of Yorkshire. Cave. Sci. 15, 15-19.

Pentecost, A. & Terry, C. 1989. Inability to demonstrate calcite precipitation by bacterial isolates from travertine. Geomicrobiol. J. 6, 185-94.

Pentecost, A. & Tortora, P. 1989. Bagni di Tivoli, Lazio: a modern travertine-depositing site and its associated microorganisms. Boll. Soc. Geol. Ital. 108, 315-324.

Pentecost, A. 1989. Calothrix-dominated oncolites from northern England. In Fifth International Symposium on Biomineralization (Ed. R. E. Crick). pp.443-54.

Pentecost, A. 1989. Observations on the Scytonema mats of San  Salvador, Bahamas. In Proceedings of the Fourth  Symposium on the Geology of the Bahamas. (ed. J.  Mylroie)
pp. 295-302.

Pentecost, A. 1989. Discovery of Phormidium stromatolites in Grahams Harbour, San Salvador. ibid pp. 303-4.

Pentecost, A., Thorpe, P.M., Harkness, D. D.& Lord, T. C. 1990. Some radiocarbon dates for tufas of the Craven district of Yorkshire. Radiocarbon, 32: 93-7.

Pentecost, A. 1990. The formation of travertine shrubs: Mammoth Hot Springs, Wyoming. Geol. Mag., 127:  159-168.

Pentecost, A. 1990. The algal flora of travertine: an  overview. In: Travertine-marl: stream deposits in  Virginia. (Eds. J. S. Herman & D. A. Hubbard).  Virginia Division of Mineral Resources Publication  101: 117-128, Charlottesville, Virginia.

Pentecost, A. 1990. Calcification processes in Algae and Cyanobacteria. In: Calcareous Algae and  Stromatolites. (Riding, R., ed.) pp. 3-20. Springer-Verlag, Berlin.

Pentecost, A. 1990.  The tractive force generated by a  motile, filamentous cyanobacterium. Microbios Lett. 44, 111-117.

Spiro, B. & Pentecost, A. 1990. The diurnal carbon balance at Waterfall Beck, Yorkshire. NERC Isotope Geosciences Laboratory Report 1987-1990 : 14-14.

Pentecost, A. & Spiro, B. 1990. Stable carbon isotope composition of calcites associated with modern freshwater cyanobacteria and algae. Geomicrobiol. J. 8, 17-26.

Pentecost, A. 1990. The identity of suspended particles in a calcite-depositing stream and their significance in trapping and binding phenomena. Limnologica. 21, 251-255.

Pentecost, A. 1991. Coccolith accumulation rates: Cretaceous to Recent. In: Mechanisms and Phylogeny  of mineralization in Biological Systems.  (Eds. S.  Suga & H. Nakahara)
pp.369-373. Springer-Verlag, Tokyo.

Pentecost, A. 1991. The weathering rates of some sandstone cliffs, central Weald, England. Earth Surface Processes & Landforms. 16, 83-91.

Pentecost, A. 1991. Algal and bryophyte flora of a Yorkshire (U.K) hill stream: a comparative approach using biovolume estimations. Arch. Hydrobiol.121, 181-201.

Pentecost, A. 1991. A note on the vegetation of some English crons. Naturalist. 116, 125-131.

Pentecost, A. 1991. A new and interesting site for the calcite-encrusted desmid, Oocardium stratum Naeg. in the British Isles. Br. phycol. J. 26, 297-301.

Pentecost, A. 1991. Springs that turn life to stone. New Scientist. 21/28 Dec. 42-44.

Pentecost, A. 1992. Travertine: life inside the rock. Biologist. 39, 161-164.

Pentecost, A. 1992. A note on the colonisation of limestone rocks by cyanobacteria. Arch. Hydrobiol. 124, 167-172.

Pentecost, A. 1992. Growth and distribution of endolithic algae in some North Yorkshire streams (UK). Br. phycol. J. 27, 145-151.

Sarfati, J. & Pentecost, A. 1992. Tussocky microstructure, a biologic event in Upper Proterozoic stromatolites: comparisons with modern freshwater stromatolite-builders. In: Early Organic Evolution (Ed. M. Schidlowski, S. Golubic, M.M. Kimberley, D. M. McKirdy & P.A. Trudinger) pp. 468-477. Springer, Berlin.

Spiro, A. & Pentecost, A. 1991. One day in the life of a stream - a diurnal carbon mass-balance and its fossil record (Waterfall Beck, Yorkshire). Geomicrobiol. J. 9, 1-11.

James, P. W. & Pentecost, A. 1992. Opegrapha. In: The lichen flora of Britain & Ireland. (Ed. by W.O. Purvis et al.) pp. 404-415. Natural History Museum, London. 

Pentecost, A. 1992. Carbonate chemistry of surface waters in a temperate karst region, the southern Yorkshire dales. [Effects of aquatic, esp. cyanobacterium photosynthesis on
carbonate chemistry & calcification] J. Hydrol. 139,  211-232.

Pentecost, A. 1993. British travertines: a review. Proc. Geol. Ass. 104, 23-39.
Pentecost, A. & Chang, M. S. 1993. Eutrophication in two Chinese Lakes. In: Quest
for a better China
(Ed. by the Forum Team of the Alliance for a better China, pp. 185-194. Global Publishing Co. Inc. USA.

Pentecost, A. 1993. Field relationships between scytonemin density, growth, and irradiance in cyanobacteria occurring in low illumination regimes. Microbial Ecology. 26, 101-11.

Goudie, A. S, Viles, H. A. & Pentecost, A. 1993. The Late-Holocene tufa decline in Europe. The Holocene 3, 181-186.

Pentecost, A. 1993. The Chalybeate Springs at Tunbridge Wells, their properties and medicinal value. 11pp. Privately  published. 

Pentecost, A. 1993. An account of the algae growing  upon the statuary at Villa Volkonsky, Rome with recommendations regarding the preservation of the sculpture. Report to the British Embassy, Rome 6 pp.

Viles, H. C. & Pentecost, A. 1994. Problems in assessing the weathering action of lichens with an example of epiliths on sandstone. In. Robinson, D. A.  & Williams, R. G. (Eds.) Rock weathering and Landform Evolution. pp. 99-116. J. Wiley, Chichester. International Geomorphological Conference, University  o f Sussex September, 1992.

Pentecost, A. 1994. Formation of laminate travertines at Bagno Vignone, Italy. Geomicrobiol. J. 12 : 239-52.
Pentecost, A. 1995. Geochemistry of carbon dioxide in six travertine-depositing waters of Italy. J. Hydrol. 167, 263-78.

Pentecost, A. & Viles, H. A. 1994. A review and reasssessment of travertine classification. Geogr. phys. et Quaternaire. 48, 305-14.

Pentecost, A. 1995. British thermophilic cyanobacteria. Arch. Hydrobiol. 132, 407-414.

Flinn, D. & Pentecost, A. 1995. Travertine-cemented screes on the serpentinite seacliffs of Unst and Fetlar, Shetland. Min. Mag. 59, 259-65.

Pentecost, A. 1995. The microbial ecology of some  Italian hot-spring travertines. Microbios. 81, 45-58.

Pentecost, A. 1995. Significance of the biomineralizing microniche in a Lyngbya (cyanobacterium) travertine. Geomicrobiol. J. 13, 213-222.

Pentecost, A. 1996. The Quaternary travertine deposits of Europe and Asia Minor. Quat. Sci Rev. 14, 1005-1028.

Pentecost, A. 1996.  Moss growth and travertine deposition: significance of photosynthesis, evaporation and degassing of carbon dioxide. J Bryol. 19, 229-34.

Jones, T.P., Fortier, S. M., Pentecost, A. & Collinson, M. A.1996. Stable carbon and oxygen isotopic compositions of Recent charophyte oosporangia and water from Malham Tarn. U.K.: palaeontological implications. Biogeochemistry. 34, 99-112.

Pentecost, A. 1996. High temperature ecosystems and chemical interactions with their environment. In: Bock, G and Goode, J (Eds). Evolution of  Hydrothermal Ecosystems onEarth (and  Mars?).  pp.99-108. Ciba Foundation Symposium 202. J. Wiley, Chichester.

Pentecost, A. & Ponsford, S. 1996. Some new observations on the mud springs of Wootton Bassett,  Wiltshire. Geology Today214-5 (Sept-Oct. issue).

Pentecost, A., Bailey, R. G., Busulwa, H. S. & Williams, A.1997. Epilithic algal communities of the Bujuku-Mubuku River System, Rwenzori Mountains, Uganda. Arch. Hydrobiol. 139, 479-493.

Pentecost, A. 1997. Lichen and moss communities on  rocky coasts. In: Ecosystems of the World: Dry Coastal Ecosystems vol. 2C (E. Van der Maarel, Ed.). pp. 195-205. Elsevier, Amsterdam.

Pentecost, A., Bayari, S. & Yesertener, C. 1997. Phototrophic organisms of the Pamukkale Travertine, Turkey: their distribution and influence on travertine deposition. Geomicrobiol J. 14, 269-83.

Pentecost, A. 1998. Phosphorus fractionation in the sediments of Malham Tarn, North Yorkshire. Fld.  Stud9, 337-42.

Pentecost, A. 1998.  Some observations on the biomass  and distribution of cryptogamic epiphyte in the Upper Montane Forest of the Rwenzori Mountains, Uganda. Global Ecology and Biogeography Lett7, 273-84.

Pentecost, A. 1998. The significance of calcite (travertine) formation by algae in a moss-dominated travertine from Matlock Bath, England.  Arch. Hydrobiol. 143, 487-509.

Pentecost, A. 1999. The origin and development of the travertines and associated thermal waters at Matlock Bath, Derbyshire. Proc. Geol. Ass. 110, 217-32.

Pentecost, A. 1999. Analysing Environmental Data. Addison-Wesley, U.K.  216 pp.

Pentecost, A. & Askew, K. 1999. The thermal springs of Matlock Bath their history and properties: with  an account of the Matlock petrifying wells. 26pp. Privately published.

Viles, H. A. & Pentecost, A. 1999. Geomorphological controls on tufa deposition at Nash Brook, South Wales, United Kingdom. Cave& Karst Science. 26, 61-8.

Pentecost, A. 1999. A note on sand ripples developing in sandstone rock seepages of the Weald, U.K. Earth Surface proc. Landforms. 24, 1257-9.

Zhang, Z. & Pentecost, A. 1999. Bryophyte communities associated with travertine formation at Yorkshire National Park, U.K. Carsalogica Sinica. 18, 366-74.

Pentecost, A., Peterken, G. F. Viles, H. C. 2000. The travertine dams of Slade Brook, Gloucestershire: their formation and conservation. Geology Today. 16, 22-25.

Pentecost, A. 2000. A note on the stable carbon isotope composition of bryophytes in calcareous aquatic habitats and its relationship to carbon dioxide assimilation. J. Bryol. 22, 13-15.

Pentecost, A. 2000. Some observations in the erosion of Tarn Moss by the waters of Malham Tarn. Field Studies 9, 569-81.

Pentecost, A. & Whitton, B. A. 2000. Cyanobacteria and Limestone. In: Whitton, B. A. & Potts, M. Eds. The Ecology of Cyanobacteria.  pp. 257-79. Kluwer, Amsterdam.

Zhang, Z. & Pentecost, A. 2000. New and noteworthy bryophytes from the travertines of Guizhou and Sichuan, S.W. China. J. Bryol. 22, 66-8.

Pentecost, A. & Peppiatt, D. 2000. A modern tree ring analysis from Tarn Moss, Malham Tarn, Yorkshire. Naturalist. 125, 167-171.

Pentecost, A. & Zhang, Z. 2000. The travertine flora of Jiuzhaigou and Munigou, China, and its relationship with calcium carbonate deposition. Cave and Karst Science. 27, 71-8.

Pentecost, A., Goudie, A., Viles, H. & Keen, D.H. 2000. The travertine deposit of Shelsley Walsh, Hereford & Worcestershire. Trans. Woolh. Nat. Fld. Club. 50, 25-36.

Pentecost, A. & Zhang, Z. 2001. A note on freshwater research in China, with some observations on the algae from Doupe Pool, Guizhou Province. Freshwater Forum. 15, 77-84.

John, D. M., Pentecost, A. & Whitton, B.A. 2001. Terrestrial and freshwater eukaryotic algae. In: Hawksworth, D.L. (Ed.) The changing wildlife of Great Britain and Ireland. Taylor & Francis, London. Syst. Assn. Special Series vol. 62, 148-50.

Seward, M. R. D. & Pentecost, A. 2001. The lichen flora of the Malham Tarn area. Fld. Stud. 10, 57-92.

Coletta, P., Pentecost, A. & Spiro, B. 2001. Stable isotopes in charophyte incrustations: relationships with climate and water chemistry. Palaeogeography, Palaeoclimatology, Palaeoecology. 173, 9-19.

Pentecost, A. & Zhang, Z. 2001. A review of Chinese Travertines. Cave & Karst Science. 28, 15-28.

Zhang, Z. & Pentecost, A. 2001. A study of the bryophyte communities of karst cave twilight zones in the Rhone Alps, France. Carsalogica Sinica. 20, 236-240.
Pentecost, A. 2002. Microdistribution of algae in an Italian thermogene travertine. Arch. Hydrobiol. 152, 439-449.

Pentecost, A. & Happey-Wood, C. M. 2002.  Freshwater Algae. In: The Plant Life of Snowdonia (edited by Rhind, P. & Evans, D.) pp. 137-148, Gomer Press, Ceridigion.

Pentecost, A. 2002. Lichens In: The Plant Life of Snowdonia (edited  by Rhind, P. & Evans, D.) pp. 118-136. Gomer Press, Ceridigion. [n.b. This book has won two awards for its content and appeal to general readers].

Pentecost, A. 2002. Raphidophyta In: The Freshwater Algae of the British Isles (Ed. John, D.M.,  Whitton, B.A. & Brook, A.J. ). pp. 208-210. Cambridge University Press.

Pentecost, A. 2002. Tetrasporales. Ibidem. pp. 299-303.

Pentecost, A. 2002. Volvocales. Ibidem. pp. 303-327.

Huxley, R. & Pentecost, A. 2002. Oedogoniales. Ibidem. pp. 409-433.

Pentecost, A. and Zhang. Z. 2002. Bryophytes from some travertine-depositing sites in France and the U.K.: relationships with climate and water chemistry. J. Bryol. 24, 233-241.

Zhang, Z. & Pentecost, A. 2002. The floristic characteristics of the bryophytes and Karst deposition in Karst Caves, England. Acta Bot. Boreal.-Occident. Sin. 22, 359-67.

Zhang, Z. & Pentecost, A. 2002. The bryophyte communities associated with travertine deposition in northwestern England and North Wales, U.K. Guihaia. 22, 45-49.

Zhang, Z. & Pentecost, A. 2002. The floristic characteristics of tufa bryophytes and the main travertine deposition types in the U.K. Carsalogica Sinica. 21, 36-43.

Pentecost, A. 2002. Analysing data. In: The Student's companion to Geography. Edited by Rogers, A. & Viles, H. A. pp. 203-211. Blackwell, Oxford.

Pentecost, A. & Williams, J. D. 2002. Some notes on the microclimate associated with the lichen Thamnolia vermicularis var. subuliformis from Snowdon, North Wales. Graphis scripta. 13, 59-63.

Baron, M., Pentecost, A. & Parnell, J. 2003. Hot and cold spring deposits of as a source of palaeo fluid samples on Mars.

Nunez, R., Spiro, B., Pentecost, A., Kim, A. & Coletta, P. 2003. Organo-geochemical and stable isotope indicators of environmental change in a marl lake, Malham Tarn, North Yorkshire, U.K. J. Paleolimnol. 28, 403-417.

Pentecost, A. & Edwards, H. G. M. 2003. Raman spectroscopy and light microscopy of a modern and sub-fossil microstromatolite: Rivularia haematites (cyanobacteria, Nostocales).  Int. J. Astrobiol. 1, 357-363.

Pentecost, A. 2003. A preliminary investigation into ascospore characters in the Graphidaceae and Opegraphaceae. B.L.S. Bull. 92, 35-39.

Pentecost, A. 2003. Taxonomic identity, ecology and distribution of the calcite-depositing cyanobacterium Phormidium incrustatum (Oscillatoriaceae). Crypt. Algol. 24, 307-321.

Pentecost, A., Jones, B. & Renaut, R. W. 2003. What is a hot spring? Can J. Earth Sci. 40, 1443-1446.

Pentecost, A. 2003. Cyanobacteria associated with hot spring travertines. Can J. Earth Sci. 40, 1447-1457.

Andrews, J., Coletta, P., Pentecost, A., Riding, R. & Spiro, B. 2004. Calcification of charophyte gyrogonites and its significance in the fossil record. Palaeogeography, Palaeoclimatology, Palaeoecology.

Pentecost, A. 2004. Travertine In: Encyclopedia of Cave and Karst science. (Ed. By J. Gunn). pp. 736-8, Fitzroy Dearborn, New York.

Pentecost, A. 2004. The travertines of Huanglong and Jiuzhaigou Scenic Areas, Sichuan. pp. 423-5. ibidem.

Pentecost, A. 2004. Cave entrance habitats. pp. 318-19. Ibidem.

Robinson, C. H., Saunders, P. W., Madan, N. J., Pyrce-Miller, E. J. & Pentecost, A. 2004. Does nitrogen deposition affect soil microfungal diversity and soil N and P dynamics in a high Arctic ecosystem? Global Change Biol. 10, 1-15.

Pentecost, A. 2004. Significance of marine algal calcification inhibitors in the global carbon cycle. Thalassas. 20, 45-49.

Clayden, S. R., Pentecost, A. & Dawson, R. J. MacG. 2004. Growth of the lichen Rhizocarpon lecanorinum, with comments on the Aplin-Hill and Lichenometric curves. Symbiosis. 37, 379-393.

Pentecost, A. & Coletta, P. (2004). A note on the travertines of Suio, Roccamonfina with reference to their microbial communities and geochemical origins. Geol. Romana. 37, 109-112.

Pentecost, A. 2005. Travertine. Springer-Verlag, Berlin and Heidelberg. 445 pp.

Pentecost, A. 2005. The freshwater-algal flora of the Kent and Sussex Weald, UK: 160 years of change. Arch. Hydrobiol. 163, 259-285.

Pentecost, A. & Zhang, Z. 2005. A note on the distribution of plants in Scoska Cave, N. Yorks, North Yorkshire, United Kingdom, and their relationship to light intensity. Cave and Karst Science. 31, 119-122.

Pentecost, A. 2005. Hot springs, thermal springs and warm springs. What's the difference? Geology Today. 21, 222-224.

Gilbert, O. L., Goldie, H., Hodgson, D., Marker, M., Pentecost, A., Proctor, M. & Richardson, D. 2005. The ecology of Cowside Beck, a tributary of the River Skirfare in the Malham area of Yorkshire. Malham Tarn Field Centre. 45pp.

Pentecost, A. 2005. Freshwater Algae of the Tunbridge Wells Area including Ashdown Forest. King's College London. 21pp.

Pentecost, A. 2006. The cryptogam-flora of Eridge Park, Sussex with some observations on forest structure and management. 26 pp. King's College London. Restricted view report.

Pentecost, A. & Zhang, Z. 2006. Response of bryophytes to exposure and water availability on some European travertines. J. Bryol. 28, 21-26.

Pentecost, A., Andrews, J. E., Dennis, P. F., Marca-Bell, A. & Dennis, S. 2006. Charophyte growth in small temperate water bodies: extreme isotopic disequilibrium an implications for the palaeoecology of shallow marl lakes. [cant find reprints at present]

Pentecost, A. 2006. A biography of the Sussex botanist Edward Jenner [1803-1827]. Queckett. J. Microsc. 40, 257-267.

Pentecost, A. & Colleta, P. 2007. The role of photosynthesis and CO2 evasion in travertine formation: a quantitative investigation at an important travertine-depositing hot spring, Le Zitelle, Lazio, Italy. J. Geol. Soc. 164, 843-853.

Zhang, Z-H., Chen, J-K. & Pentecost, A. 2007. Ecologic study on aquatic bryophyte communities  in stream karst waterfalls in the Alps, France. Carsalogica Sinica. 26, 23-30.

Viles, H. A. & Pentecost, A. 2007. Tufa and Travertine. In: Nash, D. J. & McLaren, S. J. (Eds.) Geochemical Sediments and Landscapes. pp. 173-199. Blackwell, Oxford.

Pentecost, A. & Zhang Z. H. 2008. Microfossils and geochemistry of some modern, Holocene and Pleistocene travertines from North Yorkshire and Derbyshire. Proc. Yorks. Geol Soc. 57, 79-94.

Pentecost, A. 2008. River Macrophyte Sampling: methodologies and variability. Environment Agency Report. 17 pp.

Zhang, Z-H., Chen, J-K. & Pentecost, A. 2008. Aquatic communities of bryophytes associated with karst waterfalls in England. Acta Hydrobiologica Sinica. 32, 134-140.

Pentecost, A. 2009. Remarks on the identity of the subaerial green alga Cephaleuros endophyticus (F.E. Fritsch) Printz and a new record of the species from Northern England. Naturalist. 134, 41-44.

Pentecost, A. 2009. Permanent quadrats for saxicolous lichens in Britain. BLS Bulletin. 104, 2-5.

Pentecost, A. 2009. The marl lakes of the British Isles. Freshwater Reviews. 2, 167-197.

Pentecost, A. & James, P.W.  2009. Opegrapha. In: The lichen flora of Britain & Ireland.2nd Fully revised Edition. (Ed. by W.O. Purvis et al.) pp. 404-415. Natural History Museum, London. 

Pentecost, A. 2010. Continental Carbonates- Preservation of natural and historic heritage sites. In: Carbonates in Continental Settings. Ed. By A.  M. Alonso-Zarza & L. Taner. Developments in Sedimentology 62: 297-314.

Pentecost, A. & Williamson, D. B. 2010. Some new observations on desmids in Northern England with particular reference to the genus Mesotaenium Naegeli. Queckett Journal of Microscopy. 41, 227-231.

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